Brothers in Frame: Comparing the LC250, GX550 and 4Runner

Brothers in Frame: Comparing the LC250, GX550 and 4Runner on the TNGA-F

One platform, three takes on adventure – the 4Runner, Land Cruiser 250 and Lexus GX550

In recent years Toyota has refreshed its off-road lineup in spectacular fashion, launching three closely related models: the 6th-gen Toyota 4Runner, the Toyota Land Cruiser 250 (Prado J250) and the Lexus GX550. While each of them is aimed at a slightly different audience, they share the same DNA – the modern TNGA-F platform, which underpins Toyota’s latest generation of off-road machines. These three SUVs represent different interpretations of a single idea: proven mechanical strength, serious off-road capability and a built-in readiness for adventure. Here we take a closer look at what sets them apart, what they share – and which one makes the most sense as an expedition vehicle. For that reason, we’ll focus on their more off-road-oriented trims.

Styling

Although all three share the same TNGA-F architecture, they differ not only in character but also in appearance. The GX550 and LC250 are distinguished mainly by their front ends – the Lexus goes for a massive brand-signature grille and more futuristic headlights, while the Land Cruiser opts for a simpler, more utilitarian design. In the First Edition, the LC250 came with either classic round headlights (with slightly weaker lighting performance) or rectangular Bi-LED units. At the rear the differences are mostly in the details – the GX550 features an elegant illuminated strip connecting the taillights, while the LC250 keeps things straightforward with a no-frills look.

The new 6th-gen 4Runner clearly pays homage to its predecessors – its design lines echo elements from earlier generations, something fans of the classic silhouette will surely appreciate. Its front end also borrows cues from the Toyota Tacoma, reinforcing its practical, adventure-ready character and setting it apart from the more luxurious Lexus and the more conservative Land Cruiser.

One of the key changes compared to previous generations of the GX and Land Cruiser Prado is the tailgate design – gone is the traditional side-hinged door, replaced by an upward-lifting hatch. This has both advantages (easier access when space on the sides is limited) and drawbacks (less convenient to open when carrying gear on the roof). The 4Runner, meanwhile, has used this setup for years.

All three models also feature an independently opening rear window. In the GX and Land Cruiser, the glass tilts upward manually, while the 4Runner stands out with a power-operated drop-down window – a feature especially appreciated by dog owners and anyone hauling long items without having to open the entire hatch.

Exterior Dimensions

Model	Length	Width	Height	Wheelbase
GX550	around. 500 cm	200 cm	193 cm	285 cm
LC250	around. 492 cm	198 cm	198 cm	285 cm
4Runner 6gen	around. 483 cm	193 cm	185 cm	279 cm

Dimensions may vary slightly depending on trim level.

Interior

Each of the three models – GX550, LC250 and 4Runner – takes a different approach to cabin design, reflecting its target audience.

The Lexus GX550 offers the most luxurious interior of the group. Expect high-quality materials (leather, metal accents, soft-touch plastics), a large high-resolution display, digital gauges and premium-grade audio.

The Toyota Land Cruiser 250 is the most utilitarian – the cabin is functional and solid, with plenty of physical buttons designed to withstand dirt and heavy use. Features like a 12” central screen and digital instruments are there, but everything is oriented toward usability in tough conditions.

The 6th-gen Toyota 4Runner strikes a balance between the two – its design is modern and more lifestyle-oriented, yet still focused on practicality. Durable materials, a 14” display and smart details like grab handles, power outlets and scratch-resistant finishes are all part of the package. There are even optional switches pre-wired for additional electrical accessories such as lights. The cabin sits noticeably lower than in the other two, which might be an issue for taller passengers. In the hybrid versions of both the 4Runner and LC, the floor is raised by about 12 cm, which cuts into usable space a bit. That said, the cargo area is larger than in the previous generation across all three models.

In short – the GX550 stands for comfort and elegance, the LC250 emphasizes durability and simplicity, while the 4Runner blends versatility with a modern adventure-ready vibe.

Engines

The lineup offers four engine options: a 2.4L, a 2.4L Hybrid, a 2.8L Diesel and a 3.5L V6 exclusive to the Lexus.

It’s no surprise that after test drives the most enjoyable and responsive choice is the 3.5L V6. The vehicle accelerates briskly, runs smoothly with no vibrations and, paired with a 10-speed automatic, simply glides down the road. The downside is fuel consumption – while the manufacturer claims 12.3 L/100 km, in reality it’s closer to 14 L/100 km.

The 2.4L Hybrid, available in both the LC250 (US) and the 4Runner, comes close to the Lexus in performance, while offering lower fuel use at around 12 L/100 km.

The standard 2.4L, offered in the 4Runner and LC250 (GCC), delivers sufficient power despite weaker specs, though it’s no speed demon. The lower peak torque is noticeable compared to the other engines.

The 2.8L Diesel, a proven unit carried over from previous generations but now paired with a new transmission, also provides adequate performance, though refinement is significantly worse. Toyota quotes 10.3 L/100 km, but in our experience it’s closer to 12 L/100 km (possibly down to driving style).

For comparison, the well-known 4.0L V6 from the previous Land Cruiser 150 and 4Runner produced 270 hp and 378 Nm at 4400 rpm, while the 2.4L turbo reaches peak torque as early as 1700 rpm – and that difference is very noticeable behind the wheel.

One more point for buyers to consider: in Poland hybrid vehicles benefit from lower excise tax. This means that although a 4Runner Hybrid is more expensive at the dealer, after import it actually ends up costing less than the same model with a traditional powertrain.

 

Model	Engine	Power	Torque	Transmission	Drive
GX550 Overtrail	3.4 V6 Twin Turbo	349 KM	649 Nm	10-speed automatic	4x4 with low-range transfer case
LC250 Hybrid	2.4 I4 Turbo Hybrid	326 KM	630 Nm	8-speed automatic	4x4 with low-range transfer case
LC250 Diesel	2.8 I4 Turbo Diesel	ok. 204 KM	500 Nm	8-speed automatic	4x4 with low-range transfer case
4Runner 2.4 Turbo	2.4 I4 Turbo	278 KM	430 Nm	8-speed automatic	4x4 with low-range transfer case
4Runner TRD Pro	2.4 I4 Turbo Hybrid (iForce MAX)	326 KM	630 Nm	8-speed automatic	4x4 with low-range transfer case
4Runner Trailhunter	2.4 I4 Turbo Hybrid (iForce MAX)	326 KM	630 Nm	8-speed automatic	4x4 with low-range transfer case

Drivetrain

Part-time vs. full-time

The Land Cruiser and GX550, thanks to their center differential, offer full-time four-wheel drive – a convenient and safe solution on any surface. In the 4Runner this system is reserved for the more road-focused Limited and Platinum trims. The other variants, including the off-road TRD Pro and Trailhunter, use a part-time setup, meaning the front axle must be engaged manually. As a result, 4WD should only be used on loose surfaces like mud, snow or gravel. Driving in 4WD on dry pavement is not recommended and can damage the drivetrain. In practice this demands more awareness from the driver, especially when road conditions change quickly, such as in winter.

Rear differential

The 6th-gen Toyota 4Runner and Lexus GX550 received a larger differential with a 9.5” ring gear. For reasons unknown, the LC250 uses the smaller 8.5” unit carried over from the previous Prado. Of course, this is a proven setup from the LC150 and FJ Cruiser and makes little difference for most owners running stock or only slightly larger tires.

CV joints

The Land Cruiser 250 keeps three-pronged CV joints on the front half-shafts, just like the previous generation. The 4Runner and Lexus GX550, however, are fitted with double-offset joints, which not only allow for greater articulation but are also stronger.

Off-road equipment

●       GX550 Overtrail: full-time 4WD, rear and center diff lock, e-KDSS (electronic sway bar disconnect), air suspension, 33” A/T tires, off-road cameras, crawl control, Multi-Terrain Select.

●       LC250: full-time 4WD, rear diff lock, Crawl Control, Multi-Terrain Select, off-road cameras, front sway bar disconnect.

●       4Runner Trailhunter: part-time 4WD, ARB Old Man Emu off-road suspension, 33” A/T tires, steel skid plates, snorkel, rear diff lock, Multi-Terrain Monitor, front sway bar disconnect.

●       4Runner TRD Pro: part-time 4WD, FOX suspension with remote reservoirs, 33” A/T tires, steel skid plates, rear diff lock, Multi-Terrain Select, off-road cameras, front sway bar disconnect.

Another key difference lies in the sway bar system. In the 4Runner and LC250 only the front bar can be disconnected, increasing front axle articulation. The Lexus GX550, however, uses e-KDSS, which automatically disconnects both front and rear sway bars – resulting in far greater suspension travel (better articulation) while still improving stability on the road during cornering.

Ground Clearance and Angles

Model	Approach Angle	Departure Angle	Ramp breakover	Ground Clearance
GX550 Overtrail	26°	24°	23°	approx. 220–240 mm (adjustable)
LC250	31°	22°	25°	approx. 220 mm
4Runner Trailhunter	32°	24°	24°	approx. 220 mm

Most of the specs are quite similar, except for the GX’s approach angle, where the “big jaw” front end is clearly a drawback.

Summary: Which model suits whom?

GX550 Overtrail: The right choice for those who want to combine high driving comfort with true off-road capability. With its rich equipment, advanced suspension systems and luxurious interior, the GX is perfect for long expeditions without compromise. It’s the most expensive of the three, but also offers the highest level of comfort and technology.

LC250 (Hybrid or Diesel): An ideal option for those who value simplicity, durability and proven solutions. The diesel version is better suited for remote regions where high-octane fuel may be hard to find, while the hybrid offers stronger performance and smoother operation. The LC250 is a solid foundation for adventures, priced right in the middle of the pack.

4Runner: A model aimed at fans of classic styling and off-road modifications. In factory trims like the Trailhunter and TRD Pro it already comes with a wealth of off-road gear, and thanks to massive aftermarket support it can easily be customized to fit individual needs. It’s also the most budget-friendly option of the three, delivering the best value in terms of capability per dollar.

Availability

In the official European lineup only the Land Cruiser 250 with the 2.8 Diesel is offered. That doesn’t mean the other models are out of reach – individual import is possible.

It’s worth noting that hybrid-powered vehicles benefit from a lower excise tax in Poland, making the LC250 Hybrid price comparable to the diesel version, and even allowing the Toyota 4Runner Hybrid to come in slightly cheaper.

The Lexus GX550, however, is a different story – due to its high base price and no excise tax breaks, its final cost after import and taxation will be by far the highest of the three